(
  (
    (
      Drosophila_melanogaster,
      Caenorhabditis_elegans
    )
,
    (
      (
        Ciona_intestinalis,
        Ciona_savignyi
      )
,
      (
        (
          Eptatretus_burgeri,
          Petromyzon_marinus
        )
,
        (
          (
            (
              (
                Danio_rerio,
                Astyanax_mexicanus
              )
,
              (
                (
                  (
                    (
                      (
                        Poecilia_formosa,
                        Xiphophorus_maculatus
                      )
,
                      Oryzias_latipes
                    )
,
                    Oreochromis_niloticus
                  )
,
                  (
                    (
                      Tetraodon_nigroviridis,
                      Takifugu_rubripes
                    )
,
                    Gasterosteus_aculeatus
                  )

                )
,
                Gadus_morhua
              )

            )
,
            Lepisosteus_oculatus
          )
,
          (
            (
              (
                (
                  (
                    (
                      (
                        (
                          (
                            (
                              (
                                (
                                  (
                                    (
                                      (
                                        (
                                          (
                                            (
                                              (
                                                (
                                                  Cricetulus_griseus_chok1gshd,
                                                  Cricetulus_griseus_crigri
                                                )
,
                                                Mesocricetus_auratus
                                              )
,
                                              Peromyscus_maniculatus_bairdii
                                            )
,
                                            Microtus_ochrogaster
                                          )
,
                                          (
                                            (
                                              (
                                                (
                                                  (
                                                    (
                                                      (
                                                        (
                                                          (
                                                            (
                                                              (
                                                                (
                                                                  (
                                                                    Mus_musculus,
                                                                    Mus_musculus_BALB/cJ
                                                                  )
,
                                                                  (
                                                                    (
                                                                      Mus_musculus_C3H/HeJ,
                                                                      Mus_musculus_CBA/J
                                                                    )
,
                                                                    Mus_musculus_DBA/2J
                                                                  )

                                                                )
,
                                                                Mus_musculus_AKR/J
                                                              )
,
                                                              (
                                                                (
                                                                  Mus_musculus_FVB/NJ,
                                                                  Mus_musculus_NOD/ShiLtJ
                                                                )
,
                                                                Mus_musculus_domesticus_WSB/EiJ
                                                              )

                                                            )
,
                                                            (
                                                              Mus_musculus_129S1/SvImJ,
                                                              Mus_musculus_LP/J
                                                            )

                                                          )
,
                                                          (
                                                            Mus_musculus_reference_CL57BL6,
                                                            Mus_musculus_C57BL/6NJ
                                                          )

                                                        )
,
                                                        Mus_musculus_NZO/HlLtJ
                                                      )
,
                                                      Mus_musculus_musculus_PWK/PhJ
                                                    )
,
                                                    Mus_musculus_castaneus_CAST/EiJ
                                                  )
,
                                                  Mus_spretus
                                                )
,
                                                Mus_caroli
                                              )
,
                                              Mus_pahari
                                            )
,
                                            Rattus_norvegicus
                                          )

                                        )
,
                                        Nannospalax_galili
                                      )
,
                                      Jaculus_jaculus
                                    )
,
                                    Dipodomys_ordii
                                  )
,
                                  (
                                    (
                                      (
                                        (
                                          (
                                            Cavia_porcellus,
                                            Cavia_aperea
                                          )
,
                                          Chinchilla_lanigera
                                        )
,
                                        Octodon_degus
                                      )
,
                                      (
                                        (
                                          Heterocephalus_glaber_female,
                                          Heterocephalus_glaber_male
                                        )
,
                                        Fukomys_damarensis
                                      )

                                    )
,
                                    Ictidomys_tridecemlineatus
                                  )

                                )
,
                                (
                                  Oryctolagus_cuniculus,
                                  Ochotona_princeps
                                )

                              )
,
                              Tupaia_belangeri
                            )
,
                            (
                              (
                                (
                                  (
                                    (
                                      (
                                        (
                                          (
                                            (
                                              Pan_troglodytes,
                                              Pan_paniscus
                                            )
,
                                            Homo_sapiens
                                          )
,
                                          Gorilla_gorilla
                                        )
,
                                        Pongo_abelii
                                      )
,
                                      Nomascus_leucogenys
                                    )
,
                                    (
                                      (
                                        (
                                          (
                                            (
                                              Cercocebus_atys,
                                              Mandrillus_leucophaeus
                                            )
,
                                            Papio_anubis
                                          )
,
                                          (
                                            (
                                              Macaca_fascicularis,
                                              Macaca_mulatta
                                            )
,
                                            Macaca_nemestrina
                                          )

                                        )
,
                                        Chlorocebus_sabaeus
                                      )
,
                                      (
                                        (
                                          Rhinopithecus_roxellana,
                                          Rhinopithecus_bieti
                                        )
,
                                        Colobus_angolensis_palliatus
                                      )

                                    )

                                  )
,
                                  (
                                    (
                                      (
                                        Cebus_capucinus,
                                        Saimiri_boliviensis_boliviensis
                                      )
,
                                      Callithrix_jacchus
                                    )
,
                                    Aotus_nancymaae
                                  )

                                )
,
                                Carlito_syrichta
                              )
,
                              (
                                (
                                  Microcebus_murinus,
                                  Propithecus_coquereli
                                )
,
                                Otolemur_garnettii
                              )

                            )

                          )
,
                          (
                            (
                              Erinaceus_europaeus,
                              Sorex_araneus
                            )
,
                            (
                              (
                                (
                                  (
                                    (
                                      (
                                        Sus_scrofa,
                                        Vicugna_pacos
                                      )
,
                                      (
                                        (
                                          Ovis_aries,
                                          Capra_hircus
                                        )
,
                                        Bos_taurus
                                      )

                                    )
,
                                    Tursiops_truncatus
                                  )
,
                                  Equus_caballus
                                )
,
                                (
                                  (
                                    (
                                      Ailuropoda_melanoleuca,
                                      Mustela_putorius_furo
                                    )
,
                                    Canis_familiaris
                                  )
,
                                  (
                                    (
                                      Panthera_pardus,
                                      Panthera_tigris_altaica
                                    )
,
                                    Felis_catus
                                  )

                                )

                              )
,
                              (
                                Pteropus_vampyrus,
                                Myotis_lucifugus
                              )

                            )

                          )

                        )
,
                        (
                          Dasypus_novemcinctus,
                          Choloepus_hoffmanni
                        )

                      )
,
                      (
                        (
                          Loxodonta_africana,
                          Procavia_capensis
                        )
,
                        Echinops_telfairi
                      )

                    )
,
                    (
                      (
                        Sarcophilus_harrisii,
                        Notamacropus_eugenii
                      )
,
                      Monodelphis_domestica
                    )

                  )
,
                  Ornithorhynchus_anatinus
                )
,
                (
                  (
                    (
                      (
                        (
                          Gallus_gallus,
                          Meleagris_gallopavo
                        )
,
                        Anas_platyrhynchos
                      )
,
                      (
                        Ficedula_albicollis,
                        Taeniopygia_guttata
                      )

                    )
,
                    Pelodiscus_sinensis
                  )
,
                  Anolis_carolinensis
                )

              )
,
              Xenopus_tropicalis
            )
,
            Latimeria_chalumnae
          )

        )

      )

    )

  )
,
  Saccharomyces_cerevisiae
)
;